Even-aged stands can regenerate with many thousand seedlings per hectare before the density declines to just a few hundred trees per hectare 100 years later; management practices can lead to even lower tree numbers due to quality selection and thinning. In other words, during the development of unmanaged stands, the majority of
individuals die naturally due to competition. Despite the far-reaching consequences for structural and genetic
diversity, dead wood and fuel wood accumulation, we have only limited quantitative knowledge about the
continuous mortality of trees and the wood volume loss over longer timespans. For this study, we used a unique set of 476 unmanaged, monospecific experimental plots of Norway spruce (Picea abies (L.) H. Karst.), silver fir (Abies alba Mill.), Scots pine (Pinus sylvestris L.), European larch (Larix decidua Mill.), Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco)), European beech (Fagus sylvatica L.), and oak (Quercus robur L. and Quercus petraea (Matt.) Liebl.) throughout Europe to analyze the competition-based mortality of trees and its dependency on age and site conditions. First, we show that the total stem volume production, standing stock, and mortality were continuously increasing until an age of 100–150 years. The accumulated competition-caused stem volume loss at that age amounted to 500–1000 m3 ha 1. Second, the net growth of the stands (share of the growth that is accumulated in the standing stock) strongly decreased with increasing age even when the gross growth was still high. The proportion of the net growth versus gross growth continuously decreased with increasing age regardless of site quality. Third, we show a degressive decrease of the annual relative tree number mortality rates from 0.05 to 0.20 in young down to 0.01–0.02 in mature stands. For some species, we found these rates to be site dependent with different directions of the site effect. The interplay of decreasing mortality rates and increasing average volume of the dead trees resulted in unimodal mortality curves over time of the annual mortality, peaking at 3–12 m3 ha 1 yr 1 at ages of about 75–150 years. Keywords: Competition-based mortality, Self-thinning, Natural stem volume turnover, Carbon storage, Standing stock vs. total stand production, Long-term observation
