Stems, of various sizes and shapes, are involved in most of the organic processes and interactions of plants, ranging from support, transport, and storage to development and protection. The stem itself is a crucially important intermediary: it links above - and belowground organs - connecting roots to leaves. An international team of leading researchers vividly illustrate that stems are more than pipes, more than simple connecting and supporting structures; rather stems are critical, anatomically distinct structures of enormous variability. It is, to an unappreciated extent, this variability that underpins both the diversity and the success of plants in myriad ecosystems.